Check-list and distribution maps of the molluscs of the Czech and Slovak Republics

Correspondence: horsak@sci.muni.cz

Suggested citation:
Horsák M., Čejka T., Juřičková L., Beran L., Horáčková J., Dvořák L., Coufal R., Maňas M. & Horsáková V., 2023: Check-list and distribution maps of the molluscs of the Czech and Slovak Republics. – Online at http://mollusca.sav.sk/malacology/checklist.htm, checklist updated at December 2, 2023, maps updated at April 18, 2023. https://doi.org/10.5281/zenodo.10263078

The current version of the check-list follows Horsák et al. (2013) with up-to-date modifications supplemented with a reference. Distribution map for each species was constructed based on published data and our knowledge based on unpublished records. So far 251 species of molluscs, including 223 species of gastropods (51 aquatic and 172 terrestrial) and 28 species of bivalves, have been found outdoors in the Czech Republic. The fauna of Slovakia comprises 260 species, including 231 gastropods (55 aquatic and 176 terrestrial) and 29 bivalves.

Abbreviations:
To the checklist: B – Bohemia; M – Moravia; S – Slovakia; ( ) – probably extinct species for the territory; [ ] – an extinct species for the territory; * – a non-native species for the territory; ? – an occurrence is probable, but not reliably confirmed yet; ! – an endemic species of smaller area.
To the maps: M – past distribution; R – recent distribution; M<R – number of occurrences increased over the last few decades (e.g. for non-native species); M>R – number of occurrences declined over the last few decades (e.g. for seriously threatened species); M=R – no obvious changes in the species distribution.

Updates are marked by underlining and commented below the list of species.

GASTROPODA
subclass Neritimorpha
order Cycloneritida
Neritidae
Theodoxus Montfort, 1810
T. danubialis (C. Pfeiffer, 1828)	–	M	S	map	images
T. fluviatilis (Linnaeus, 1758)	[B]	–	S*	map	images
T. transversalis (C. Pfeiffer, 1828)	–	–	S	map	images
subclass Caenogastropoda
order Architaenioglossa
Viviparidae
Viviparus Montfort, 1810
V. acerosus Bourguignat, 1862	B*	M	S	map	images
V. contectus (Millet, 1813)	B	M	S	map	images
V. viviparus (Linnaeus, 1758)	B	M*	–	map	images
Aciculidae
Acicula Hartmann, 1821
A. parcelineata (Clessin, 1911)	–	M	S	map	images
Platyla Moquin-Tandon, 1856
P. polita (Hartmann, 1840)	B	M	S		images
unassigned order of Caenogastropoda
Melanopsidae
Esperiana Bourguignat, 1877
E. daudebartii acicularis (A. Férussac, 1823)	–	–	S	map	images
E. esperi (A. Férussac, 1823)	–	–	S	map	images
order Littorinimorpha
Bithyniidae
Bithynia Leach, 1818
B. leachii (Sheppard, 1823)	–	M	S	map	images
B. tentaculata (Linnaeus, 1758)	B	M	S	map	images
B. transsilvanica (Bielz, 1853)	B*	M	S	map	images
Bythinellidae
Bythinella Moquin-Tandon, 1856
B. austriaca (von Frauenfeld, 1857) – see comment	B	M	S	map	images
B. pannonica (von Frauenfeld, 1865)	–	–	!S	map	images
Hydrobiidae
Clathrocaspia Lindholm, 1930
C. knipowitschii (Makarov, 1938)	–	–	S*	map	images
Alzoniella Giusti & Bodon, 1984
A. slovenica (Ložek & Brtek, 1964)	–	!M	!S	map	images
Hauffenia Pollonera, 1889
Hauffenia kissdalmae Erőss & Petró, 2008	–	–	!S	map	images
Hauffenia lozekiana Haase, Grego, Eröss, Farkas & Fehér, 2021	–	–	!S	map	images
Lithoglyphidae
Lithoglyphus Menke, 1830
L. naticoides (C. Pfeiffer, 1828)	–	M	S	map	images
Tateidae
Potamopyrgus Stimpson, 1865
P. antipodarum (Gray, 1843)	B*	M*	S*	map	images
subclass Heterobranchia
infraclass "Lower Heterobranchia"
Valvatidae
Valvata O. F. Müller, 1773
V. cristata O. F. Müller, 1774	B	M	S	map	images
V. macrostoma Mörch, 1864	B	–	S	map	images
V. piscinalis (O. F. Müller, 1774)	B	M	S	map	images
Borysthenia Lindholm, 1914
B. naticina (Menke, 1845)	–	–	S	map	images
superorder Hygrophila
Acroloxidae
Acroloxus Beck, 1838
A. lacustris (Linnaeus, 1758)	B	M	S	map	images
Lymnaeidae
Galba Schrank, 1803
G. truncatula (O. F. Müller, 1774)	B	M	S	map	images
Stagnicola Jeffreys, 1830
S. corvus (Gmelin, 1791)	B	M	S	map	images
S. fuscus (C. Pfeiffer, 1821)	B	–	–	map	images
S. palustris (O. F. Müller, 1774)	B	–	–	map	images
S. turricula (Held, 1836)	B	M	S	map	images
Ladislavella Dybowski, 1913
L. occulta (Jackiewicz, 1959)	B	M	S	map	images
Radix Montfort, 1810
R. ampla (Hartmann, 1821)	B	M	S	map	images
R. auricularia (Linnaeus, 1758)	B	M	S	map	images
R. balthica (Linnaeus, 1758)	B	M	S	map	images
R. labiata (Rossmässler, 1835)	B	M	S	map	images
R. lagotis (Schrank, 1803)	B	M	S	map	images
Myxas J. Sowerby, 1822
M. glutinosa (O. F. Müller, 1774)	[B]	–	–	map	images
Lymnaea Lamarck, 1799
L. stagnalis (Linnaeus, 1758)	B	M	S	map	images
Physidae
Aplexa Fleming, 1820
A. hypnorum (Linnaeus, 1758)	B	M	S	map	images
Physa Draparnaud, 1801
P. acuta Draparnaud, 1805	B*	M*	S*	map	images
P. fontinalis (Linnaeus, 1758)	B	M	S	map	images
Planorbidae
Planorbis O. F. Müller, 1773
P. carinatus O. F. Müller, 1774	B	M	S	map	images
P. planorbis (Linnaeus, 1758)	B	M	S	map	images
Anisus Studer, 1820
A. leucostoma (Millet, 1813)	B	M	S	map	images
A. septemgyratus (Rossmässler, 1835)	–	M	S	map	images
A. spirorbis (Linnaeus, 1758)	B	M	S	map	images
A. vortex (Linnaeus, 1758)	B	M	S	map	images
A. vorticulus (Troschel, 1834)	B	M	S	map	images
Bathyomphalus Charpentier, 1837
B. contortus (Linnaeus, 1758)	B	M	S	map	images
Gyraulus Charpentier, 1837
G. acronicus (A. Férussac, 1807)	B	M	S	map	images
G. albus (O. F. Müller, 1774)	B	M	S	map	images
G. crista (Linnaeus, 1758)	B	M	S	map	images
G. parvus (Say, 1817)	B	M	S	map	images
G. riparius (Westerlund, 1865)	–	–	S	map	images
G. rossmaessleri (Schmidt, 1852)	B	M	S	map	images
Hippeutis Charpentier, 1837
H. complanatus (Linnaeus, 1758)	B	M	S	map	images
Segmentina Fleming, 1818
S. nitida (O. F. Müller, 1774)	B	M	S	map	images
Ancylus O. F. Müller, 1773
A. fluviatilis O. F. Müller, 1774	B	M	S	map	images
Planorbarius Duméril, 1806
P. corneus (Linnaeus, 1758)	B	M	S	map	images
Menetus H. & A. Adams, 1855
M. dilatatus (Gould, 1841)	B*	–	–	map	images
Ferrissia Walker, 1903
F. californica (Rowell, 1863)	B*	M*	S*	map	images
superorder Eupulmonata
order Ellobiida
Ellobiidae
Carychium O. F. Müller, 1773
C. minimum O. F. Müller, 1774	B	M	S		images
C. tridentatum (Risso, 1826)	B	M	S		images
order Stylommatophora
Succineidae
Succinea Draparnaud, 1801
S. putris (Linnaeus, 1758)	B	M	S		images
Oxyloma Westerlund, 1885
O. elegans (Risso, 1826)	B	M	S		images
Succinella Mabille, 1871
S. oblonga (Draparnaud, 1801)	B	M	S		images
Quickella C. Boettger, 1939
Q. arenaria (Potiez & Michaud, 1838)	–	–	S	map	images
Chondrinidae
Abida Turton, 1831
A. secale (Draparnaud, 1801)	–	–	S	map	images
Granaria Held, 1838
G. frumentum (Draparnaud, 1801)	B	M	S		images
Chondrina Reichenbach, 1828
C. avenacea (Bruguière, 1792)	B	–	–		images
C. arcadica clienta (Westerlund, 1883)	B	M	S	map	images
C. tatrica Ložek, 1948	–	–	!S	map	images
Truncatellinidae
Columella Westerlund, 1878
C. aspera Waldén, 1966	B	M	S	map	images
C. columella (G. von Martens, 1830)	–	–	S	map	images
C. edentula (Draparnaud, 1805)	B	M	S		images
Truncatellina Lowe, 1852
T. claustralis (Gredler, 1856)	B	M	S	map	images
T. costulata (Nilsson, 1823)	B	M	S	map	images
T. cylindrica (A. Férussac, 1807)	B	M	S		images
Argnidae
Argna Cossmann, 1889
A. bielzi (Rossmässler, 1859)	–	–	S	map	images
Cochlicopidae
Cochlicopa A. Férussac, 1821
C. lubrica (O. F. Müller, 1774)	B	M	S		images
C. lubricella (Porro, 1838)	B	M	S		images
C. nitens (M. von Gallenstein, 1848)	B	M	S	map	images
Enidae
Chondrula Beck, 1837
C. tridens (O. F. Müller, 1774)	B	M	S		images
Ena Turton, 1831
E. montana (Draparnaud, 1801)	B	M	S		images
Merdigera Held, 1838
M. obscura (O. F. Müller, 1774)	B	M	S		images
Zebrina Held, 1838
Z. detrita (O. F. Müller, 1774)	B	M	S	map	images
Orculidae
Orcula Held, 1838
O. dolium (Draparnaud, 1801)	–	M	S	map	images
Sphyradium Charpentier, 1837
S. doliolum (Bruguière, 1792)	B	M	S		images
Pagodulinidae
Pagodulina Clessin, 1876
P. pagodula (Des Moulins, 1830)	–	M	S	map	images
Pupillidae
Pupilla Fleming, 1828
P. alpicola (Charpentier, 1837)	B	M	S	map	images
P. muscorum (Linnaeus, 1758)	B	M	S		images
P. sterrii (Forster, 1840)	B	M	S		images
P. triplicata (Studer, 1820)	B	M	S		images
Pyramidulidae
Pyramidula Fitzinger, 1833
P. pusilla Gittenberger & Bank, 1996	B	M	S	map	images
P. saxatilis (Hartmann, 1842)	–	–	S	map	images
Spelaeodiscidae
Spelaeodiscus Brusina, 1886
S. triarius tatricus (Hazay, 1883)	–	–	!S	map	images
Valloniidae
Vallonia Risso, 1826
V. costata (O. F. Müller, 1774)	B	M	S		images
V. declivis Sterki, 1893	B	–	S	map	images
V. enniensis (Gredler, 1856)	B	M	S	map	images
V. excentrica Sterki, 1893	B	M	S		images
V. pulchella (O. F. Müller, 1774)	B	M	S		images
Acanthinula Beck, 1847
A. aculeata (O. F. Müller, 1774)	B	M	S		images
Vertiginidae
Vertigo O. F. Müller, 1773
V. alpestris Alder, 1838	B	M	S		images
V. angustior Jeffreys, 1830	B	M	S	map	images
V. antivertigo (Draparnaud, 1801)	B	M	S		images
V. geyeri Lindholm, 1925	B	M	S	map	images
V. lilljeborgi (Westerlund, 1871)	B	M	–	map	images
V. arctica (Wallenberg, 1858)	–	–	S	map	images
V. moulinsiana (Dupuy, 1849)	B	M	S	map	images
V. pusilla O. F. Müller, 1774	B	M	S		images
V. pygmaea (Draparnaud, 1801)	B	M	S		images
V. ronnebyensis (Westerlund, 1871)	B	–	–	map	images
V. substriata (Jeffreys, 1833)	B	M	S		images
Clausiliidae
Alopia H. & A. Adams, 1855
A. bielzi clathrata (E. A. Bielz, 1856)	–	–	!S	map	images
Cochlodina A. Férussac, 1821
C. cerata (Rossmässler, 1836)	–	!M	S	map	images
C. costata (C. Pfeiffer, 1828)	B	M	–		images
C. dubiosa corcontica Brabenec, 1967	!B	–	–		images
C. fimbriata (Rossmässler, 1835)	–	–	S	map	images
C. laminata (Montagu, 1803)	B	M	S		images
C. orthostoma (Menke, 1828)	B	M	S		images
Charpentieria Stabile, 1864
C. ornata (Rossmässler, 1836)	B	M	–		images
Ruthenica Lindholm, 1924
R. filograna (Rossmässler, 1836)	B	M	S		images
Pseudofusulus H. Nordsieck, 1977
P. varians (C. Pfeiffer, 1828)	B	[M]	S	map	images
Macrogastra Hartmann, 1841
M. badia (C. Pfeiffer, 1828)	B	M	–	map	images
M. borealis (Boettger, 1878)	–	M	S	map	images
M. plicatula (Draparnaud, 1801)	B	M	S		images
M. tumida (Rossmässler, 1835)	B	M	S		images
M. ventricosa (Draparnaud, 1801)	B	M	S		images
Clausilia Draparnaud, 1805
C. bidentata (Ström, 1765)	B	–	–	map	images
C. cruciata (Studer, 1820)	B	M	S		images
C. dubia Draparnaud, 1805	B	M	S		images
C. pumila C. Pfeiffer, 1828	B	M	S		images
C. rugosa (Draparnaud, 1801)	B	M	S		images
Laciniaria Hartmann, 1842
L. plicata (Draparnaud, 1801)	B	M	S		images
Alinda H. & A. Adams, 1855
A. biplicata (Montagu, 1803)	B	M	S		images
Balea Gray, 1824
B. perversa (Linnaeus, 1758)	B	M	S		images
Pseudalinda Boettger, 1877
P. stabilis (L. Pfeiffer, 1847)	–	–	S	map	images
Vestia Hesse, 1916
V. elata (Rossmässler, 1836)	–	–	S		images
V. gulo (E. A. Bielz, 1859)	–	M	S	map	images
V. ranojevici moravica (Brabenec, 1952)	–	!M	–	map	images
V. turgida (Rossmässler, 1836)	B	M	S		images
Bulgarica O. Boettger, 1877
B. cana (Held, 1836)	B	M	S		images
B. nitidosa (Uličný, 1893)	!B	–	–		images
Ferussaciidae
Cecilioides A. Férussac, 1814
C. acicula (O. F. Müller, 1774)	B	M	S		images
C. petitiana (Benoit, 1862)	–	–	S*	map	images
Punctidae
Punctum Morse, 1864
P. pygmaeum (Draparnaud, 1801)	B	M	S	map	images
Helicodiscidae
Lucilla R. T. Lowe, 1852
L. scintilla (R. T. Lowe, 1852)	B*	?M*	S*	map	images
L. singleyana (Pilsbry, 1889)	?B*	?M*	S*	map	images
Discidae
Discus Fitzinger, 1833
D. perspectivus (Megerle von Mühlfeld, 1816)	B	M	S		images
D. rotundatus (O. F. Müller, 1774)	B	M	S		images
D. ruderatus (Hartmann, 1821)	B	M	S		images
Gastrodontidae
Aegopinella Lindholm, 1927
A. epipedostoma iuncta Hudec, 1964	–	M	S		images
A. minor (Stabile, 1864)	B	M	S		images
A. nitens (Michaud, 1831)	B	M	S		images
A. nitidula (Draparnaud, 1805)	B	M	–		images
A. pura (Alder, 1830)	B	M	S		images
A. ressmanni (Westerlund, 1883)	B	–	–	map	images
Nesovitrea Cooke, 1921
N. hammonis (Ström, 1765)	B	M	S		images
N. petronella (L. Pfeiffer, 1853)	B	M	S		images
Zonitoides Lehmann, 1862
Z. arboreus (Say, 1817)	–	M*	–	map	images
Z. nitidus (O. F. Müller, 1774)	B	M	S		images
Oxychilidae
Oxychilus Fitzinger, 1833
O. alliarius (Miller, 1822)	B	–	–	map	images
O. cellarius (O. F. Müller, 1774)	B	M	S		images
O. depressus (Sterki, 1880)	B	M	S		images
O. deubeli (A. J. Wagner, 1914)	–	–	S	map	images
O. draparnaudi (Beck, 1837)	B*	M*	S*		images
O. glaber (Rossmässler, 1835)	B	M	S		images
O. hydatinus (Rossmässler, 1838)	–	–	S*	map	images
O. inopinatus (Uličný, 1887)	B	M	S		images
O. mortilleti (L. Pfeiffer, 1859)	(B)	–	–	map	images
Carpathica A. J. Wagner, 1895
C. calophana (Westerlund, 1881)	–	–	S	map	images
Daudebardia Hartmann, 1821
D. brevipes (Draparnaud, 1805)	B	M	S		images
D. rufa (Draparnaud, 1805)	B	M	S	map	images
Pristilomatidae
Vitrea Fitzinger, 1833
V. contracta (Westerlund, 1871)	B	M	S		images
V. crystallina (O. F. Müller, 1774)	B	M	S		images
V. diaphana (Studer, 1820)	B	M	S		images
V. subrimata (Reinhardt, 1871)	B	M	S		images
V. transsylvanica (Clessin, 1877)	B	M	S	map	images
Euconulidae
Euconulus Reinhardt, 1883
E. alderi (Gray, 1840)	B	M	S		images
E. fulvus (O. F. Müller, 1774)	B	M	S		images
Zonitidae
Aegopis Fitzinger, 1833
A. verticillus (Lamarck, 1822)	B	M	–		images
Milacidae
Tandonia Lessona & Pollonera, 1882
T. budapestensis (Hazay, 1880)	B	M	S	map	images
T. kusceri (Wagner, 1931)	–	–	S*	map	images
T. rustica (Millet, 1843)	B	–	–	map	images
Agriolimacidae
Deroceras Rafinesque, 1820
D. agreste (Linnaeus, 1758)	B	M	S	map	images
D. invadens Reise, Hutchinson, Schunack & Schlitt, 2011	B*	M*	S*	map	images
D. juranum Wüthrich, 1993	B	–	–	map	images
D. laeve (O. F. Müller, 1774)	B	M	S	map	images
D. praecox Wiktor, 1966	B	M	S	map	images
D. reticulatum (O. F. Müller, 1774)	B	M	S	map	images
D. rodnae Grossu & Lupu, 1965	B	M	S	map	images
D. sturanyi (Simroth, 1894)	B	M	S	map	images
D. turcicum (Simroth, 1894)	B	M	S	map	images
Krynickillus Kaleniczenko, 1851
K. melanocephalus (Kaleniczenko, 1851)	–	–	S*	map	images
Boettgerillidae
Boettgerilla Simroth, 1910
B. pallens Simroth, 1912	B*	M*	S*		images
Limacidae
Ambigolimax Pollonera, 1887
A. valentianus (A. Férussac, 1821)	–	–	S*	map	images
Bielzia Clessin, 1887
B. coerulans (M. Bielz, 1851)	–	M	S	map	images
Limax Linnaeus, 1758
L. cinereoniger Wolf, 1803	B	M	S		images
L. maximus Linnaeus, 1758	B	M	S		images
Limacus Lehmann, 1864
L. flavus (Linnaeus, 1758)	B*	M*	S	map	images
L. maculatus (Kaleniczenko, 1851)	–	M*	–	map	images
Malacolimax Malm, 1868
M. tenellus (O. F. Müller, 1774)	B	M	S		images
Lehmannia Heynemann, 1863
L. carpatica Hutchinson, Reise & Schlitt, 2022	–	M	S	map	images
L. macroflagellata Grossu & Lupu, 1962	B	M	S	map	images
L. marginata (O. F. Müller, 1774)	B	M	S		images
Vitrinidae
Semilimax Gray, 1847
S. kotulae (Westerlund, 1883)	B	M	S		images
S. semilimax (J. Férussac, 1802)	B	M	S		images
Eucobresia H. B. Baker, 1929
E. diaphana (Draparnaud, 1805)	B	M	S		images
E. nivalis (Dumont & Mortillet, 1854)	B	M	S		images
Vitrina Draparnaud, 1801
V. pellucida (O. F. Müller, 1774)	B	M	S		images
Arionidae
Arion A. Férussac, 1819
A. circumscriptus Johnston, 1828	B	M	?S		images
A. distinctus Mabille, 1868	B	M	S		images
A. fasciatus (Nilsson, 1823)	B	M	S		images
A. fuscus (O. F. Müller, 1774)	B	M	S		images
A. intermedius Normand, 1852	B	M	S*	map	images
A. obesoductus Reischütz, 1973	B	M	S*	map	images
A. rufus (Linnaeus, 1758)	B	M	S		images
A. silvaticus Lohmander, 1937	B	M	S		images
A. vulgaris Moquin-Tandon, 1855	B*	M*	S*		images
Camaenidae
Fruticicola Held, 1838
F. fruticum (O. F. Müller, 1774)	B	M	S		images
Helicodontidae
Helicodonta A. Férussac, 1821
H. obvoluta (O. F. Müller, 1774)	B	M	S		images
Hygromiidae
Euomphalia Westerlund, 1889
E. strigella (Draparnaud, 1801)	B	M	S		images
Monacha Fitzinger, 1833
M. cartusiana (O. F. Müller, 1774)	B*	M	S		images
M. claustralis (Rossmässler, 1834)	B*	–	–	map	images
Trochulus Chemnitz, 1786
T. hispidus (Linnaeus, 1758)	B	M	S		images
T. striolatus danubialis (Clessin, 1874)	–	–	S	map	images
T. villosulus (Rossmässler, 1838)	–	M	S		images
Petasina Beck, 1847
P. bakowskii (Poliński, 1924)	–	–	S	map	images
P. bielzi (E. A. Bielz, 1860)	–	–	S	map	images
P. edentula (Draparnaud, 1805)	B	–	–	map	images
P. filicina (L. Pfeiffer, 1841)	–	–	S	map
P. unidentata (Draparnaud, 1805)	B	M	S		images
Plicuteria Schileyko, 1978
P. lubomirski (Ślósarski, 1881)	B	M	S		images
Perforatella Schlüter, 1838
P. bidentata (Gmelin, 1791)	B	M	S		images
P. dibotrion (E. A. Bielz, 1860)	–	–	S	map	images
Pseudotrichia Schileyko, 1970
P. rubiginosa (Rossmässler, 1838)	B	M	S		images
Monachoides Gude & Woodward, 1921
M. incarnatus (O. F. Müller, 1774)	B	M	S		images
M. vicinus (Rossmässler, 1842)	B	M	S		images
Urticicola Lindholm, 1927
U. umbrosus (C. Pfeiffer, 1828)	B	M	S		images
Lozekia Hudec, 1970
L. transsilvanica (Westerlund, 1876)	–	–	S	map	images
Hygromia Risso, 1826
H. cinctella (Draparnaud, 1801)	B*	–	S*	map	images
Geomitridae
Helicopsis Fitzinger, 1833
H. striata (O. F. Müller, 1774)	(B)	[M]	S	map	images
Candidula Kobelt, 1871
C. unifasciata (Poiret, 1801)	(B)	M	S		images
Helicella A. Férussac, 1821
H. itala (Linnaeus, 1758)	B	M	–	map	images
Xerolenta Monterosato, 1892
X. obvia (Menke, 1828)	B	M	S		images
Cernuella Schlüter, 1838
C. neglecta (Draparnaud, 1805)	B*	–	–		images
Helicidae
Arianta Turton, 1831
A. arbustorum (Linnaeus, 1758)	B	M	S		images
Helicigona A. Férussac, 1821
H. lapicida (Linnaeus, 1758)	B	M	–		images
Faustina Kobelt, 1904
F. cingulella (Rossmässler, 1837)	–	–	S	map	images
F. faustina (Rossmässler, 1835)	B	M	S		images
F. rossmaessleri (L. Pfeiffer, 1848)	–	–	S	map	images
Isognomostoma Fitzinger, 1833
I. isognomostomos (Schröter, 1784)	B	M	S		images
Causa Schileyko, 1971
C. holosericea (Studer, 1820)	B	M	S		images
Cepaea Held, 1838
C. hortensis (O. F. Müller, 1774)	B	M	S		images
C. nemoralis (Linnaeus, 1758)	B*	M*	S*	map	images
Caucasotachea Boettger, 1909
C. vindobonensis (C. Pfeiffer, 1828)	B	M	S	map	images
Cornu Born, 1778
C. aspersum (O. F. Müller, 1774)	B*	M*	S*	map	images
Helix Linnaeus, 1758
H. lucorum Linnaeus, 1758	B*	–	S*	map	images
H. lutescens Rossmässler, 1837	–	–	S	map	images
H. pomatia Linnaeus, 1758	B	M	S		images
H. thessalica Boettger, 1886	–	M	S	map	images
BIVALVIA	occurrence
Heteroconchia
Palaeoheterodonta
Unionoida
Margaritiferidae
Margaritifera Schumacher, 1815
M. margaritifera (Linnaeus, 1758)	B	[M]	–	map	images
Unionidae
Unio Philipsson, 1788
U. crassus Philipsson, 1788	B	M	S	map	images
U. pictorum (Linnaeus, 1758)	B	M	S	map	images
U. tumidus Philipsson, 1788	B	M	S	map	images
Anodonta Lamarck, 1799
A. anatina (Linnaeus, 1758)	B	M	S	map	images
A. cygnea (Linnaeus, 1758)	B	M	S	map	images
Pseudanodonta Bourguignat, 1877
P. complanata (Rossmässler, 1835)	B	M	S	map	images
Sinanodonta Modell, 1945
S. woodiana (Lea, 1834)	B*	M*	S*	map	images
Heterodonta
Veneroida
Corbiculidae
Corbicula Megerle von Mühlfeld, 1811
C. fluminea (O. F. Müller, 1774)	B*	M*	S*	map	images
Sphaeriidae
Sphaerium Scopoli, 1777
S. corneum (Linnaeus, 1758)	B	M	S		images
S. lacustre (O. F. Müller, 1774)	B	M	S		images
S. nucleus (Studer, 1820)	B	M	S		images
S. rivicola (Lamarck, 1818)	B	M	S		images
S. solidum (Normand, 1844)	–	–	S*	map	images
Pisidium C. Pfeiffer, 1821
P. amnicum (O. F. Müller, 1774)	B	M	S		images
Euglesa Jenyns, 1832
E. casertana (Poli, 1791)	B	M	S		images
E. globularis (Clessin, 1873)	B	M	S		images
E. henslowana (Sheppard, 1825)	B	M	S		images
E. hibernica (Westerlund, 1894)	B	M	S		images
E. milium (Held, 1836)	B	M	S		images
E. nitida (Jenyns, 1832)	B	M	S		images
E. obtusalis (Lamarck, 1818)	B	M	S		images
E. personata (Malm, 1855)	B	M	S		images
E. pseudosphaerium (J. Favre, 1927)	B	M	S		images
E. subtruncata (Malm, 1855)	B	M	S		images
E. supina (A. Schmidt, 1851)	B	M	S		images
Odhneripisidium Kuiper, 1962
O. moitessierianum (Paladilhe, 1866)	B	M	S		images
O. tenuilineatum (Stelfox, 1918)	B	M	S		images
Dreissenidae
Dreissena Van Beneden, 1835
D. polymorpha (Pallas, 1771)	B*	M*	S*		images
D. rostriformis (Deshayes, 1838)	–	–	S*	map	images

Comment to Bythinella species

Bythinella austriaca (von Frauenfeld, 1857) – There are three more species, by their shell morphology similar to B. austriaca, reported from Slovakia: B. hungarica (Hazay, 1881), B. metarubra (Falniowski, 1987) and B. steffeki Grego & Glöer, 2019. The latter species has been described based on a single population in the Slovenský kras Mts., the area with frequent occurrence of B. austriaca, using only shell and genital morphology. It has been unshakeably proven (e.g. Bichain et al. 2007) that classical morphological descriptors may not constitute valid specific criteria within Bythinella. Without an integrative taxonomical study, delimiting these putative species based on both mitochondrial and nuclear gene fragments, and without the association of these species with shell morphology, we cannot accept these names as biologically valid taxa. There is no evidence to support an assumption that these species are not only extremes within (eco-)morphological plasticity of a single species, i.e. B. austriaca. Grego & Glöer (2019) stated: “The COI sequence in the GenBank under number MK673142 indicates a close relation of the new species to some of the B. austriaca clades.”, which in other words says that there is no genetic difference (at least based on COI gene) between B. austriaca and B. steffeki. The phylogenetic analysis published by Benke et al. (2009) clearly showed that B. austriaca and B. metarubra are genetically the same.

Comments to species names and new records

Viviparus acerosus Bourguignat, 1862 – for the first time recorded in Bohemia in 2015, see Beran et al. (2019).

Clathrocaspia knipowitschii (Makarov, 1938) – for the first time recorded in Slovakia in 2019, see Szekeres et al. (2022).

Hauffenia kissdalmae Erőss & Petró, 2008 – see Haase et al. (2021).

Hauffenia lozekiana Haase, Grego, Eröss, Farkas & Fehér, 2021 – see Haase et al. (2021).

Stagnicola palustris (O. F. Müller, 1774) and Stagnicola turricula (Held, 1836) – taxonomic status follows Pieńkowska et al. (2015a).

Ladislavella Dybowski, 1913 – originally as Catascopia Meier-Brook & Bargues, 2002. Nomenclature follows Pieńkowska et al. (2015a) and Pieńkowska & Lesicki (2018).

Ladislavella occulta (Jackiewicz, 1959) – European populations were for some time reported under the name L. terebra (Westerlund, 1885), see Pieńkowska & Lesicki (2018). The first record in Slovakia near the village of Senné (Čejka et al. 2020).

Radix lagotis (Schrank, 1803) – for the first records, see Čejka et al. (2021).

Physa acuta (Draparnaud, 1805) – see comments in Welter-Schultes (2012), p. 56.

Gyraulus parvus (Say, 1817) – current genetic investigation revealed that G. parvus and G. laevis are in fact part of the same species-level clade, with the former having nomenclatural priority (Lorencová et al. 2021). However, the structure within the mitochondrial tree suggests a North American origin of the invasive populations, earlier reported as G. parvus. It also suggests that although the native race in Europe, i.e. G. laevis, tend to possess some differences in conchology and ecology, the degree of overlap between the races makes it impossible to accurately distinguish between them without the DNA barcode data.

Ferrissia californica (Rowell, 1863) – originally as F. fragilis (Tryon, 1863), see Christensen (2016). Sometimes even a month matters in description of a new species.

Pupilla alpicola (Charpentier, 1837) – conspecific with Pupilla pratensis (Clessin, 1871), see Nekola et al. (2015).

Pyramidulla saxatilis (Hartmann, 1842) – species resurrected based on DNA from synonymy with P. rupestris, but it cannot be differentiated by shell morphology from P. pusilla. Only one site has been reported from Veľká Fatra Mts. based on DNA analysis of specimens from that site, see Kirchner et al. (2016) and Razkin et al. (2016, 2017).

Vertigo arctica (Wallenberg, 1858) – Nekola et al. (2018) consider it as a separate species, phylogenetically not closely related to V. modesta (Say, 1824).

Macrogastra borealis (Boettger, 1878) – Due to some nomenclatorial issues (see http://www.animalbase.uni-goettingen.de/zooweb/servlet/AnimalBase/home/speciestaxon?id=15626) the well-known name Macrogastra latestriata (Schmidt, 1857) had to be substituted by M. borealis (Boettger, 1878).

Euconulus alderi (Gray, 1840) [syn. E. praticola (Reinhardt, 1883)] – taxonomy and nomenclature follow Horsáková et al. (2020).

Tandonia kusceri (Wagner, 1931) and T. rustica (Millet, 1843) – see Korábek et al. (2016a).

Ambigolimax valentianus (Férussac, 1822) – for the first time recorded outdoors in Slovakia in 2020, see Čejka et al. (2021). Originally as Lehmannia Heynemann, 1863, for the current genus assignment see Nitz (2013) and rowson et al. (2014).

Limacus maculatus (Kaleniczenko, 1851) – for the first time recorded in Moravia in 2019, see Čejka et al. (2020).

Lehmannia carpatica Hutchinson, Reise & Schlitt, 2022 – originally reported as L. nyctelia (Bourguignat, 1861) and later also as Ambigolimax nyctelius (Bourguignat, 1861); see Nitz (2013) and rowson et al. (2014) for details. Hutchinson et al (2022) noted that the name L. nyctelia (Bourguignat, 1861) was used for five different species and raised the new name for the Carpathian populations as L. carpatica, since the original name Limax nyctelius, based on the Algerian matrerial, is now to be known as Letourneuxia nyctelia (Bourguignat, 1861).

Deroceras invadens Reise, Hutchinson, Schunack & Schlitt, 2011 – for the first time recorded in Bohemia in 2014, see Hutchinson et al. (2014); and in Slovakia in 2017, see Čejka (2018).

Deroceras juranum Wüthrich, 1993 – see Hutchinson & Reise (2009).

Krynickillus melanocephalus Kaleniczenko, 1851 – for the first time recorded outdoors in Slovakia in 2020, see Čejka et al. (2021).

Arion intermedius Normand, 1852 – for the first time recorded outdoors in Slovakia in 2020, see Čejka et al. (2021).

Trochulus hispidus (Linnaeus, 1758) – T. sericeus (Draparnaud, 1801) was confirmed to be a part of phenotypic and genetic plasticity, now being recognized as a younger synonym of T. hispidus, see Proćków et al. (2017).

Monacha claustralis (Rossmässler, 1834) – originally reported as Monacha cantiana (Montagu, 1803) by Hlaváč & Peltanová (2010); for further details see Pieńkowska et al. (2015b).

Hygromia cinctella (Draparnaud, 1801) – for the first time recorded in Slovakia in 2015, see Čejka (2015a) and Krumpálová & Holienková (2018).

Cepaea nemoralis (Linnaeus, 1758) – for the first time recorded in Slovakia in 2015, see Čejka (2015a) and Krumpálová & Holienková (2018).

Caucasotachea vindobonensis (C. Pfeiffer, 1828) – see Neiber et al. (2016).

Cornu aspersum (O. F. Müller, 1774) – for the first time recorded in Slovakia in 2015, see Čejka (2015b). For the first time recorded in Moravia in 2020, see Čejka et al. (2021).

Helix thessalica Boettger, 1886 – see Korábek et al. (2016b).

Corbicula fluminea (O. F. Müller, 1774) – for the first time recorded in Moravia in 2018, see Komzák et al. (2018).

Sphaerium lacustre (O. F. Müller, 1774) – see Bespalaya et al. (2023).

Euglesa casertana (Poli, 1791) – see Bespalaya et al. (2023).

Euglesa globularis (Clessin, 1873) – see Bespalaya et al. (2023).

Euglesa henslowana (Sheppard, 1825) – see Bespalaya et al. (2023).

Euglesa hibernica (Westerlund, 1894) – see Bespalaya et al. (2023).

Euglesa milium (Held, 1836) – see Bespalaya et al. (2023).

Euglesa nitida (Jenyns, 1832) – see Bespalaya et al. (2023).

Euglesa obtusalis (Lamarck, 1818) – see Bespalaya et al. (2023).

Euglesa personata (Malm, 1855) – see Bespalaya et al. (2023).

Euglesa pseudosphaerium (J. Favre, 1927) – the authorship should be ascribed to Favre, see Bank (2011). See Bespalaya et al. (2023).

Euglesa subtruncata (Malm, 1855) – see Bespalaya et al. (2023).

Euglesa supina (A. Schmidt, 1851) – see Bespalaya et al. (2023).

Odhneripisidium moitessierianum (Paladilhe, 1866) – see Bespalaya et al. (2023).

Odhneripisidium tenuilineatum (Stelfox, 1918) – see Bespalaya et al. (2023).

Dreissena rostriformis (Deshayes, 1838) – for the first time recorded in Slovakia in 2013 (Čejka et al. 2020).

References

Bank R. A., 2011: Authorships and Publication Dates in Malacology: some notes on the 2011 French Checklist of Welter-Schultes & al. – Mitteilungen der Deutschen Malakozoologischen Gesellschaft, 86: 13–24.

Benke M., Brändle M., Albrecht C. & Wilke T., 2009: Pleistocene phylogeography and phylogenetic concordance in cold‐adapted spring snails (Bythinella spp.). – Molecular Ecology, 18: 890–903. https://doi.org/10.1111/j.1365-294x.2008.04073.x

Bespalaya Y. V., Vinarski M. V., Aksenova O. V., Babushkin E. S., Gofarov M. Y., Kondakov A. V., Konopleva E. S., Kropotin A. V., Mabrouki Y., Ovchankova N. B., Palatov D. M., Sokolova S. E., Shevchenko A. R., Travina O. V., Taybi A. F., Soboleva A. A., Zubrii N. A. & Bolotov I. N., 2023: Phylogeny, taxonomy, and biogeography of the Sphaeriinae (Bivalvia: Sphaeriidae). – Zoological Journal of the Linnean Society, zlad139. https://doi.org/10.1093/zoolinnean/zlad139

Beran L., Horsák M. & Hofman S., 2019: First records of Viviparus acerosus (Bourguignat, 1862) (Gastropoda: Viviparidae) from the Czech Republic outside its native range. – Folia Malacologica, 27(3): 223–229. https://doi.org/10.12657/folmal.027.021

Bichain J. M., Gaubert P., Samadi S. & Boisselier-Dubayle M. C., 2007: A gleam in the dark: Phylogenetic species delimitation in the confusing spring-snail genus Bythinella Moquin-Tandon, 1856 (Gastropoda: Rissooidea: Amnicolidae). – Molecular Phylogenetics and Evolution, 45(3): 927–941. https://doi.org/10.1016/j.ympev.2007.07.018

Čejka T., 2015a: Three new mollusc species for Slovakia - result of one visit of the garden centre. – Malacological Bulletin,  August 7, 2015, http://malbull.blogspot.com/2015/08/four-new-mollusc-species-for-slovakia.html

Čejka T., 2015b: Brown garden snail - Cornu aspersum (O. F. Müller, 1774) in Slovakia. – Malacological Bulletin, August 4, 2015, http://malbull.blogspot.sk/2015/08/first-record-of-cornu-aspersum-o-f.html

Čejka T., 2018: First outdoor records of the slug Deroceras invadens from the Slovak Republic. – Malacological Bulletin, February 22, 2018, https://malbull.blogspot.cz/2018/02/first-outdoor-records-of-slug-deroceras.html

Čejka T., Beran L., Coufal R., Dvořák L., Hlaváč J. Č., Horáčková J., Horsáková V., Juřičková L., Kosová T., Čačaný J., Szabóová D., Říhová D., Tej B. & Horsák M., 2021: Malacological news from the Czech and Slovak Republics in 2020. – Malacologica Bohemoslovaca, 20: 56–74. https://doi.org/10.5817/MaB2021-20-56

Čejka T., Beran L., Korábek O., Hlaváč J. Č., Horáčková J., Coufal R., Drvotová M., Maňas M., Horsáková V. & Horsák M., 2020: Malacological news from the Czech and Slovak Republics in 2015–2019. – Malacologica Bohemoslovaca, 19: 71–106. https://doi.org/10.5817/MaB2020-19-71

Christensen C. C., 2016: Change of status and name for a Hawaiian freshwater limpet: Ancylus sharpi Sykes, 1900 is the invasive North American Ferrissia californica (Rowell, 1863), formerly known as Ferrissia fragilis (Tryon, 1863) (Gastropoda: Planorbidae: Ancylinae). – Bishop Museum Occasional Papers, 118: 5–8.

Grego J. & Glöer P., 2019: A new Bythinella (Mollusca: Gastropoda: Rissooidea) in the Slovak Republic. – Ecologica Montenegrina, 21: 80–85. https://doi.org/10.37828/em.2019.21.9

Haase M., Grego J., Erőss Z. P., Farkas R. & Fehér Z., 2021: On the origin and diversification of the stygobiotic freshwater snail genus Hauffenia (Caenogastropoda: Hydrobiidae) with special focus on the northern species and the description of two new species. – European Journal of Taxonomy, 775: 143–184. https://doi.org/10.5852/ejt.2021.775.1555

Hlaváč J. Č. & Peltanová A., 2010: First occurrence of the Kentish Snail Monacha cantiana (Mollusca: Gastropoda: Hygromiidae) in the Czech Republic. – Malacologica Bohemoslovaca, 9: 11–15. https://doi.org/10.5817/MaB2010-9-11

Horsák M., Juřičková L. & Picka J., 2013: Měkkýši České a Slovenské republiky. Molluscs of the Czech and Slovak Republics. – Kabourek, Zlín, 264 pp. ISBN 978-80-86447-15-5 (in Czech and English)

Horsáková V., Nekola J. C. & Horsák M., 2020: Integrative taxonomic consideration of the Holarctic Euconulus fulvus group of land snails (Gastropoda, Stylommatophora). – Systematics and Biodiversity, 18(2): 142–160. https://doi.org/10.1080/14772000.2020.1725172

Hutchinson J. M. C. & Reise H., 2009: Mating behaviour clarifies the taxonomy of slug species defined by genital anatomy: the Deroceras rodnae complex in the Sächsische Schweiz and elsewhere. – Mollusca, 27: 183–200.

Hutchinson J. M. C., Reise H. & Robinson D. G., 2014: A biography of an invasive terrestrial slug: the spread, distribution and habitat of Deroceras invadens. – Neobiota, 23: 17–64. https://doi.org/10.3897/neobiota.23.7745

Hutchinson J. M. C., Reise H. & Schlitt B., 2022: Will the real Limax nyctelius please step forward: Lehmannia, Ambigolimax, or Malacolimax? No, Letourneuxia! – Archiv für Molluskenkunde, 151(1): 19–41. https://doi.org/10.1127/arch.moll/151/019-041

Kirchner S., Harl J., Kruckenhauser L., Duda M., Sattmann H. & Haring E., 2016: Phylogeography and systematics of Pyramidula (Pulmonata: Pyramidulidae) in the eastern Alps: still a taxonomic challenge. – Journal of Molluscan Studies, 82: 110–121. https://doi.org/10.1093/mollus/eyv047

Korábek O., Čejka T. & Juřičková L., 2016a: Tandonia kusceri (Pulmonata: Milacidae), a slug new for Slovakia. – Malacologica Bohemoslovaca, 15: 3–8. https://doi.org/10.5817/MaB2016-15-3

Korábek O., Juřičková J. & Petrusek A., 2016b: Splitting the Roman snail Helix pomatia Linnaeus, 1758 (Stylommatophora: Helicidae) into two: redescription of the forgotten Helix thessalica Boettger, 1886. – Journal of Molluscan Studies, 82(1): 11–22. https://doi.org/10.1093/mollus/eyv048

Komzák P., Beran L. & Horsák M., 2018: The first record of Corbicula fluminea (O. F. Müller, 1774) in Moravia (SE Czech Republic). – Malacologica Bohemoslovaca, 17: 28–30. https://doi.org/10.5817/MaB2018-17-28

Krumpálová Z. & Holienková B., 2018: Land snails in the Slovak open-air garden centres. – Ekológia, Bratislava, 37(4): 369–379. https://doi.org/10.2478/eko-2018-0028

Lorencová E., Beran L., Nováková M., Horsáková V., Rowson B., Hlaváč J. Č., Nekola J. C. & Horsák M., 2021: Invasion at the population level: a story of the freshwater snails Gyraulus parvus and G. laevis. – Hydrobiologia, (2021) –. https://doi.org/10.1007/s10750-021-04668-w

Neiber M. T., Sagorny C. & Hausdorf B., 2016: Increasing the number of molecular markers resolves the phylogenetic relationship of ‘Cepaea’ vindobonensis (Pfeiffer 1828) with Caucasotachea Boettger 1909 (Gastropoda: Pulmonata: Helicidae). – Journal of Zoological Systematics and Evolutionary Research, 54(1): 40–45. https://doi.org/10.1111/jzs.12116

Nekola J. C., Chiba S., Coles B. F., Drost C. A., von Proschwitz T. & Horsák M., 2018: A phylogenetic overview of the genus Vertigo O. F. Müller, 1773 (Gastropoda: Pulmonata: Pupillidae: Vertigininae). – Malacologia, 62: 21–161. https://doi.org/10.4002/040.062.0104

Nekola J. C., Coles F. B. & Horsák M., 2015: Species assignment in Pupilla (Gastropoda: Pulmonata: Pupillidae): integration of DNA-sequence data and conchology. – Journal of Molluscan Studies, 81(2): 196–216. https://doi.org/10.1093/mollus/eyu083

Nitz B., 2013: Integrative systematics and biogeography of Limax (Gastropoda: Stylommatophora). – Dissertation Thesis, Ludwig-Maximilians-Universität, München, 133 pp.

Pieńkowska J. R. & Lesicki A., 2018: A note on the status of Galba occulta Jackiewicz, 1959 (Gastropoda: Hygrophila: Lymnaeidae). – Folia Malacologica, 26(4): 231–247. https://doi.org/10.12657/folmal.026.029

Pieńkowska J. R., Rybska E., Banasiak J., Wesołowska M. & Lesicki A., 2015a: Taxonomic status of Stagnicola palustris (O. F. Müller, 1774) and S. turricula (Held, 1836) (Gastropoda: Pulmonata: Lymnaeidae) in view of new molecular and chorological data. – Folia Malacologica, 23(1): 3–18. https://doi.org/10.12657/folmal.023.003

Pieńkowska J. R, Manganelli G., Giusti F. & Lesicki A., 2015b: Monacha claustralis (Rossmässler 1834) new to Polish and Czech malacofauna (Gastropoda: Pulmonata: Hygromiidae). – Journal of Conchology, 42: 79–93.

Proćków M., Kuźnik-Kowalska E., Mackiewicz P. 2017: Phenotypic plasticity can explain evolution of sympatric polymorphism in the hairy snail Trochulus hispidus (Linnaeus, 1758). – Current Zoology, 63(4): 389–402. https://doi.org/10.1093/cz/zow082

Razkin O., Sonet G., Breugelmans K., Madeira M. J., Gómez-Moliner B. J. & Backeljau T., 2016: Species limits, interspecific hybridization and phylogeny in the cryptic land snail complex Pyramidula: the power of RADseq data. – Molecular phylogenetics and evolution, 101: 267–278. https://doi.org/10.1016/j.ympev.2016.05.002

Razkin O., Gómez‐Moliner B. J., Vardinoyannis K., Martínez‐Ortí A. & Madeira M. J., 2017: Species delimitation for cryptic species complexes: case study of Pyramidula (Gastropoda, Pulmonata). – Zoologica Scripta, 46(1): 55–72. https://doi.org/10.1111/zsc.12192

Rowson B., Anderson R., Turner J. A. & Symondson W. O. C., 2014: The slugs of Britain and Ireland: undetected and undescribed species increase a well-studied, economically important fauna by more than 20%. – PLoS One. 9(3): e91907. https://doi.org/10.1371/journal.pone.0091907

Szekeres J., Beermann A., Neubauer T. A., Očadlik M., Paunović M., Raković M., Csányi B., Varga A., Weigand A., Wilke T. & Fehér Z., 2022: Rapid spread of a new alien and potentially invasive species, Clathrocaspia knipowitschii (Makarov, 1938) (Gastropoda: Hydrobiidae), in the Danube River. – Archives of Biological Sciences, 6 pp. https://doi.org/10.2298/ABS220211006S

Welter-Schultes F. W., 2012: European non-marine molluscs, a guide for species identification. – Planet Poster Editions, Göttingen, pp. A1-A3, 1-679, Q1-Q78. ISBN 978-3-933922-75-5.